Scolia (wasp)

Scolia
	Scolia hirta
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Hymenoptera
Family:	Scoliidae
Tribe:	Scoliini
Genus:	Scolia; Fabricius, 1775
Type species
	Scolia sexmaculata; (Müller, 1766)

Scolia is a genus of scoliid wasps in the subfamily Scoliinae. There are over 200 described species in Scolia.^[1]

Taxonomy and phylogeny

Scolia was described by Johan Christian Fabricius in 1775. This genus is classified under the tribe Scoliini of the subfamily Scoliinae within the family Scoliidae.^[2] Scolia is itself divided into three subgenera, Scolia (Scolia), and Scolia (Discolia), and Scolia (Hesperoscolia).^[3] Males of the subgenus Scolia (Scolia) have dense, silky setae on the volsella, while those of the subgenus Scolia (Discolia) lack dense setae on the volsella.^[2]

Description and identification

Scolia are small to medium wasps between 5–25 millimetres (0.20–0.98 in).^[2] The forewing has a single recurrent vein and two submarginal cells.^[4] The species are usually black with variable yellow or red markings. The majority of setae are usually black or white but may also be mixed with red or yellow.^[2]

Distribution

Scolia is a cosmopolitan genus. 40 species are known from India alone.^[2] Another 50 species are known from the Palearctic.^[1] 20 more species are known from the Americas.^[5]

Species

These species belong to the genus Scolia:^[6]

Scolia affinis Guérin-Méneville, 1830
Scolia anatoliae Osten, 2004
Scolia asiella Betrem, 1935
Scolia bicincta Fabricius, 1775 – double-banded scoliid
Scolia binotata Fabricius 1804
Scolia bnun Tsuneki, 1972
Scolia carbonaria (Linnaeus, 1767)
Scolia clypeata Sickmann 1894
Scolia consors (Saussure, 1863)
Scolia cypria Saussure, 1854
Scolia dubia (Say, 1837) – two-spotted scoliid wasp, blue-winged scoliid wasp
Scolia ebenina Saussure, 1858
Scolia erythrocephala Fabricius, 1798
Scolia erythropyga Burmeister, 1853
Scolia fallax Eversmann, 1849
Scolia fasciatopunctata Guerin 1838
Scolia flaviceps Eversmann, 1846
Scolia fuciformis Scopoli, 1786
Scolia galbula (Pallas, 1771)
Scolia guttata (Burmeister, 1853)
Scolia hirta (Schrank, 1781)
Scolia histrionica (Fabricius 1787)
Scolia hortorum Fabricius, 1787
Scolia kuroiwae Matsumura & Uchida 1926
Scolia mexicana (Saussure, 1858)
Scolia nigrita Fabricius
Scolia nobilis Saussure 1858
Scolia nobilitata (Fabricius, 1805) – noble scoliid wasp
Scolia oculata Matsumura 1911 – yellow-banded scoliid wasp
Scolia orientalis Saussure, 1856
Scolia picteti Saussure 1854
Scolia rufiventris (Fabricius 1804)
Scolia rugifrons Betrem 1928
Scolia semitacta Bradley, 1959
Scolia sexmaculata (O. F. Müller, 1766)
Scolia sinensis Saussure 1864
Scolia superciliaris Saussure 1864
Scolia turkestanica Betrem 1935
Scolia verticalis (Fabricius, 1775)– yellow-headed flower wasp, yellow-headed scoliid wasp
Scolia vollenhoveni Saussure 1859
Scolia wahlbergii Saussure, 1859
Scolia watanabei Matsumura 1912
† Scolia distincta Zhang, 1989
† Scolia saussureana Heer, 1865

Gallery

S. affinis in Kerala, India.
S anatoliae in Turkey.
S. bicincta female in North Carolina.
S. bilunata in Kerala, India.
S. carbonaria collected in Algeria.
S. chrysotricha in South Africa.
S. cyanipennis in Kerala, India.
S. dubia dubia in Maryland.
S. dubia haematodes in Texas.
S. fallax in Ukraine.
S. fasciatopunctata in India.
S. fuciformis in Ukraine.
S. hirta in Italy.
S. hortorum in Spain.
S. nobilitata notilitata in Virginia.
S. nobilitata otomita in Arizona.
S. nobilitata tricincta male in Colorado.
S. ocula in Japan.
S. pictei in India.
s. rufiventris in Argentina.
S. sexmaculata in Poland.
S. verticalis in Australia.
S. wahlbergii in South Africa.

References

^ ^a ^b Mokrousov, M.V.; Lelej, A.S. (2017). "62. Family Scoliidae". In Belokobylskij, S.A.; Lelej, A.S. (eds.). Annotated Catalogue of the Hymenoptera of Russia. Vol. 1. Symphyta and Apocrita: Aculeata. Saint Petersburg: Zoological Institute RAS. pp. 147–148. ISBN 978-5-98092-062-3.
^ ^a ^b ^c ^d ^e Gupta, S.K.; Jonathan, J.K. (2003). Fauna of India and the Adjacent Countries, Hymenoptera: Scoliidae. Kolkata, India: Zoological Survey of India. pp. 151–152. ISBN 81-8171-009-6.
^ Golfetti, Ivan Fernande (2019). Análise Filogenética de Scolia Fabricius (Hymenoptera, Scoliidae, Scoliinae) (MSc thesis) (in Portuguese). São Paulo State University. pp. 1–69. Retrieved 2023-07-17.
^ Augul, Razzaq Shalan (2016). "Taxonomic study of the family Scoliidae (Hymenoptera: Aculeata) in Iraq". Journal of Biodiversity and Environmental Sciences. 8 (2): 124–138. Retrieved 2023-07-17.
^ Añino, Yostin J.; Cambra, Roberto A.; Windsow, Donald M.; Zuñiga, Ronald; Quintero, Diomedes (2019). "Review of Scolia (Hymenoptera: Scoliidae) from Central America, including seasonal flight activity in Scolia guttata". Acta Biológica Colombiana. 25 (2): 202–209. doi:10.15446/abc.v25n2.77590.
^ Osten, T. (2005). "Checkliste der Dolchwespen der Welt (Insecta: Hymenoptera, Scoliidae). Teil 1: Proscoliinae und Scoliinae: Campsomerini. Teil 2: Scoliinae: Scoliini. Teil 3: Literatur" [Checklist of the Scoliidae of the World. Part 1: Proscoliinae and Scoliinae: Campsomerini. Part 2: Scoliinae: Scoliini. Part 3: Literature] (PDF). Bericht der Naturforschenden Gesellschaft Augsburg (in German). 62 (220–221): 1–62. Retrieved 2014-06-24.

[Russian-1] Mokrousov, M.V.; Lelej, A.S. (2017). "62. Family Scoliidae". In Belokobylskij, S.A.; Lelej, A.S. (eds.). Annotated Catalogue of the Hymenoptera of Russia. Vol. 1. Symphyta and Apocrita: Aculeata. Saint Petersburg: Zoological Institute RAS. pp. 147–148. ISBN 978-5-98092-062-3.

[Gupta_&_Jonathan_2003-2] Gupta, S.K.; Jonathan, J.K. (2003). Fauna of India and the Adjacent Countries, Hymenoptera: Scoliidae. Kolkata, India: Zoological Survey of India. pp. 151–152. ISBN 81-8171-009-6.

[Golfetti_2019-3] Golfetti, Ivan Fernande (2019). Análise Filogenética de Scolia Fabricius (Hymenoptera, Scoliidae, Scoliinae) (MSc thesis) (in Portuguese). São Paulo State University. pp. 1–69. Retrieved 2023-07-17.

[Augul_2016-4] Augul, Razzaq Shalan (2016). "Taxonomic study of the family Scoliidae (Hymenoptera: Aculeata) in Iraq". Journal of Biodiversity and Environmental Sciences. 8 (2): 124–138. Retrieved 2023-07-17.

[Añino_et_al._2019-5] Añino, Yostin J.; Cambra, Roberto A.; Windsow, Donald M.; Zuñiga, Ronald; Quintero, Diomedes (2019). "Review of Scolia (Hymenoptera: Scoliidae) from Central America, including seasonal flight activity in Scolia guttata". Acta Biológica Colombiana. 25 (2): 202–209. doi:10.15446/abc.v25n2.77590.

[Osten-6] Osten, T. (2005). "Checkliste der Dolchwespen der Welt (Insecta: Hymenoptera, Scoliidae). Teil 1: Proscoliinae und Scoliinae: Campsomerini. Teil 2: Scoliinae: Scoliini. Teil 3: Literatur" [Checklist of the Scoliidae of the World. Part 1: Proscoliinae and Scoliinae: Campsomerini. Part 2: Scoliinae: Scoliini. Part 3: Literature] (PDF). Bericht der Naturforschenden Gesellschaft Augsburg (in German). 62 (220–221): 1–62. Retrieved 2014-06-24.

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